The Wildcard search feature allows you to search for a specific search term(s) that appears anywhere in a name, or title when choosing a "list item" such as:
Department (Class, Project)
Any other list
When the Wildcard feature is enabled, it will allow you to search at the beginning, middle, or end of the field no matter where the words you are searching for is located within the field.
For example, if your account is named: "15001: Prepaid expenses":
If you do NOT have the wildcard turned on, then you would need to type in "15001" or "Prepaid" to find the account. Typing in "1500" would find all accounts that BEGIN with "1500", including 15001: Prepaid expenses
If you turned wildcard on, then you could type in "expenses" and it would find ANY account with the word "expenses" ANYWHERE in the name. If you typed in "prepaid" then again it would find ANY account with the word "prepaid" ANYWHERE in the name.